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Systematic studies1-4 have revealed hundreds of ultra-compact dwarf galaxies (UCDs5) in the nearby Universe. With half-light radii rh of approximately 10-100 parsecs and stellar masses M* ≈ 106-108 solar masses, UCDs are among the densest known stellar systems6. Although similar in appearance to massive globular clusters7, the detection of extended stellar envelopes4,8,9, complex star formation histories10, elevated mass-to-light ratio11,12 and supermassive black holes13-16 suggest that some UCDs are remnant nuclear star clusters17 of tidally stripped dwarf galaxies18,19, or even ancient compact galaxies20. However, only a few objects have been found in the transient stage of tidal stripping21,22, and this assumed evolutionary path19 has never been fully traced by observations. Here we show that 106 galaxies in the Virgo cluster have morphologies that are intermediate between normal, nucleated dwarf galaxies and single-component UCDs, revealing a continuum that fully maps this morphological transition and fills the `size gap' between star clusters and galaxies. Their spatial distribution and redder colour are also consistent with stripped satellite galaxies on their first few pericentric passages around massive galaxies23. The `ultra-diffuse' tidal features around several of these galaxies directly show how UCDs are forming through tidal stripping and that this evolutionary path can include an early phase as a nucleated ultra-diffuse galaxy24,25. These UCDs represent substantial visible fossil remnants of ancient dwarf galaxies in galaxy clusters, and more low-mass remnants probably remain to be found. 

Abstract We present a study of the stellar populations of globular clusters (GCs) in the Virgo Cluster core with a homogeneous spectroscopic catalog of 692 GCs within a major-axis distance R maj = 840 kpc from M87. We investigate radial and azimuthal variations in the mean age, total metallicity, [Fe/H], and α -element abundance of blue (metal-poor) and red (metal-rich) GCs using their co-added spectra. We find that the blue GCs have a steep radial gradient in [Z/H] within R maj = 165 kpc, with roughly equal contributions from [Fe/H] and [ α /Fe], and flat gradients beyond. By contrast, the red GCs show a much shallower gradient in [Z/H], which is entirely driven by [Fe/H]. We use GC-tagged Illustris simulations to demonstrate an accretion scenario where more massive satellites (with more metal- and α -rich GCs) sink further into the central galaxy than less massive ones, and where the gradient flattening occurs because of the low GC occupation fraction of low-mass dwarfs disrupted at larger distances. The dense environment around M87 may also cause the steep [ α /Fe] gradient of the blue GCs, mirroring what is seen in the dwarf galaxy population. The progenitors of red GCs have a narrower mass range than those of blue GCs, which makes their gradients shallower. We also explore spatial inhomogeneity in GC abundances, finding that the red GCs to the northwest of M87 are slightly more metal-rich. Future observations of GC stellar population gradients will be useful diagnostics of halo merger histories.